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The Jamestown settlement in the Colony of Virginia was the first permanent English settlement in the Americas. It was located on the northeast bank of the James River, about 2.5 mi (4 km) southwest of present-day Williamsburg. It was established by the London Company as "James Fort" on May 4, 1607 O.S. (May 14, 1607 N.S.), and considered permanent, after brief abandonment in 1610. It followed failed attempts, including the Roanoke Colony, established in 1585. Despite the dispatch of more supplies, only 60 of the original 214 settlers survived the 1609–1610 Starving Time. In mid-1610, the survivors abandoned Jamestown, though they returned after meeting a resupply convoy in the James River.

Jamestown served as the colonial capital from 1616 until 1699. In August 1619, the first recorded slaves from Africa to British North America arrived at present-day Old Point Comfort, near the Jamestown colony, on a British privateer ship flying a Dutch flag. The approximately 20 Africans from present-day Angola had been removed by the British crew from a Portuguese slave ship. They most likely worked in the tobacco fields, under a system of race-based indentured servitude. The modern conception of slavery in the British colonies was formalized in 1640, and fully entrenched in Virginia by 1660.

In 1676, Jamestown was deliberately burned during Bacon's Rebellion, though it was rebuilt. In 1699, the colonial capital was moved to present-day Williamsburg, Virginia. In the 18th century, Jamestown ceased to exist as a settlement and remains as an archaeological site, Jamestown Rediscovery, which houses museums and historical sites, including the Jamestown Settlement and the American Revolution Museum in Yorktown. Jamestown is one of three locations composing the Historic Triangle of Colonial Virginia, along with Williamsburg and Yorktown. Historic Jamestowne is the archaeological site on Jamestown Island and is a cooperative effort by Jamestown National Historic Site and Preservation Virginia. Jamestown Settlement, a living history interpretive site, is operated by the Jamestown Yorktown Foundation, a state agency of the Commonwealth of Virginia.

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